Cardisoma crassum, Gecarcinidae, mouthless land crab, marine invertebrates, crustaceans, decapods, Ostional Wildlife Refuge, Costa Rica, Pacific Ocean.

Remarks: Photography under controlled conditions. The mouthless land crab lives in the mangroves, where it is very difficult to photograph. To make this photo, the crab was captured and relerased into its habitat later.

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